The Structure and Distribution of Coral Reefs · Charles Darwin
We have as yet only considered the origin of barrier-reefs and atolls in their simplest form; but there remain some peculiarities in structure and some special cases, described in the two first chapters, to be accounted for by our theory. These consist--in the inclined ledge terminated by a wall, and sometimes succeeded by a second ledge with a wall, round the shores of certain lagoons and lagoon-channels; a structure which cannot, as I endeavoured to show, be explained by the simple growing powers of the corals,--in the ring or basin-like forms of the central reefs, as well as of the separate marginal portions of the northern Maldiva atolls,--in the submerged condition of the whole, or of parts of certain barrier and atoll-formed reefs; where only a part is submerged, this being generally to leeward,--in the apparent progressive disseverment of some of the Maldiva atolls,--in the existence of irregularly formed atolls, some being tied together by linear reefs, and others with spurs projecting from them,--and, lastly, in the structure and origin of the Great Chagos bank.
Step-formed ledges round certain lagoons.--If we suppose an atoll to subside at an extremely slow rate, it is difficult to follow out the complex results. The living corals would grow up on the outer margin; and likewise probably in the gullies and deeper parts of the bare surface of the annular reef; the water would encroach on the islets, but the accumulation of fresh detritus might possibly prevent their entire submergence. After a subsidence of this very slow nature, the surface of the annular reef sloping gently into the lagoon, would probably become united with the irregular reefs and banks of sand, which line the shores of most lagoons. Should, however, the atoll be carried down by a more rapid movement, the whole surface of the annular reef, where there was a foundation of solid matter, would be favourably circumstanced for the fresh growth of coral; but as the corals grew upwards on its exterior margin, and the waves broke heavily on this part, the increase of the massive polypifers on the inner side would be checked from the want of water. Consequently, the exterior parts would first reach the surface, and the new annular reef thus formed on the old one, would have its summit inclined inwards, and be terminated by a subaqueous wall, formed by the upward growth of the coral (before being much checked), from the inner edge of the solid parts of the old reef. The inner portion of the new reef, from not having grown to the surface, would be covered by the waters of the lagoon. Should a subsidence of the same kind be repeated, the corals would again grow up in a wall, from all the solid parts of the resunken reef, and, therefore, not from within the sandy shores of the lagoon; and the inner part of the new annular reef would, from being as before checked in its upward growth, be of less height than the exterior parts, and therefore would not reach the surface of the lagoon. In this case the shores of the lagoon would be surrounded by two inclined ledges, one beneath the other, and both abruptly terminated by subaqueous cliffs.
The ring or basin-formed reefs of the northern Maldiva atolls.--I may first observe, that the reefs within the lagoons of atolls and within lagoon-channels, would, if favourably circumstanced, grow upwards during subsidence in the same manner as the annular rim; and, therefore, we might expect that such lagoon-reefs, when not surrounded and buried by an accumulation of sediment more rapid than the rate of subsidence, would rise abruptly from a greater depth than that at which the efficient polypifers can flourish: we see this well exemplified in the small abruptly-sided reefs, with which the deep lagoons of the Chagos and Southern Maldiva atolls are studded. With respect to the ring or basin-formed reefs of the Northern Maldiva atolls, it is evident, from the perfectly continuous series which exists that the marginal rings, although wider than the exterior or bounding reef of ordinary atolls, are only modified portions of such a reef; it is also evident that the central rings, although wider than the knolls or reefs which commonly occur in lagoons, occupy their place. The ring-like structure has been shown to be contingent on the breaches into the lagoon being broad and numerous, so that all the reefs which are bathed by the waters of the lagoon are placed under nearly the same conditions with the outer coast of an atoll standing in the open sea. Hence the exterior and living margins of these reefs must have been favourably circumstanced for growing outwards, and increasing beyond the usual breadth; and they must likewise have been favourably circumstanced for growing vigorously upwards, during the subsiding movements, to which by our theory the whole archipelago has been subjected; and subsidence with this upward growth of the margins would convert the central space of each little reef into a small lagoon. This, however, could only take place with those reefs, which had increased to a breadth sufficient to prevent their central spaces from being almost immediately filled up with the sand and detritus driven inwards from all sides: hence it is that few reefs, which are less than half a mile in diameter, even in the atolls where the basin-like structure is most strikingly exhibited, include lagoons. This remark, I may add, applies to all coral-reefs wherever found. The basin-formed reefs of the Maldiva Archipelago may, in fact, be briefly described, as small atolls formed during subsidence over the separate portions of large and broken atolls, in the same manner as these latter were formed over the barrier-reefs, which encircled the islands of a large archipelago now wholly submerged.
Submerged and Dead Reefs.--In the second section of the first chapter, I have shown that there are in the neighbourhood of atolls, some deeply submerged banks, with level surfaces; that there are others, less deeply but yet wholly submerged, having all the characters of perfect atolls, but consisting merely of dead coral-rock; that there are barrier-reefs and atolls with merely a portion of their reef, generally on the leeward side, submerged; and that such portions either retain their perfect outline, or they appear to be quite effaced, their former place being marked only by a bank, conforming in outline with that part of the reef which remains perfect. These several cases are, I believe, intimately related together, and can be explained by the same means. There, perhaps, exist some submerged reefs, covered with living coral and growing upwards, but to these I do not here refer.
As we see that in those parts of the ocean, where coral-reefs are most abundant, one island is fringed and another neighbouring one is not fringed; as we see in the same archipelago, that all the reefs are more perfect in one part of it than in another, for instance, in the southern half compared with the northern half of the Maldiva Archipelago, and likewise on the outer coasts compared with the inner coasts of the atolls in this same group, which are placed in a double row; as we know that the existence of the innumerable polypifers forming a reef, depends on their sustenance, and that they are preyed on by other organic beings; and, lastly, as we know that some inorganic causes are highly injurious to the growth of coral, it cannot be expected that during the round of change to which earth, air, and water are exposed, the reef-building polypifers should keep alive for perpetuity in any one place; and still less can this be expected, during the progressive subsidences, perhaps at some periods more rapid than at others, to which by our theory these reefs and islands have been subjected and are liable. It is, then, not improbable that the corals should sometimes perish either on the whole or on part of a reef; if on part, the dead portion, after a small amount of subsidence, would still retain its proper outline and position beneath the water. After a more prolonged subsidence, it would probably form, owing to the accumulation of sediment, only the margin of a flat bank, marking the limits of the former lagoon. Such dead portions of reef would generally lie on the leeward side, for the impure water and fine sediment would more easily flow out from the lagoon over this side of the reef, where the force of the breakers is less than to windward; and therefore the corals would be less vigorous on this side, and be less able to resist any destroying agent. It is likewise owing to this same cause, that reefs are more frequently breached to leeward by narrow channels, serving as by ship-channels, than to windward. If the corals perished entirely, or on the greater part of the circumference of an atoll, an atoll-shaped bank of dead rock, more or less entirely submerged, would be produced; and further subsidence, together with the accumulation of sediment, would often obliterate its atoll-like structure, and leave only a bank with a level surface.
In the Chagos group of atolls, within an area of 160 miles by 60, there are two atoll-formed banks of dead rock (besides another very imperfect one), entirely submerged; a third, with merely two or three very small pieces of living reef rising to the surface; and a fourth, namely, Peros Banhos (Plate I., Figure 9), with a portion nine miles in length dead and submerged. As by our theory this area has subsided, and as there is nothing improbable in the death, either from changes in the state of the surrounding sea or from the subsidence being great or sudden, of the corals on the whole, or on portions of some of the atolls, the case of the Chagos group presents no difficulty. So far indeed are any of the above-mentioned cases of submerged reefs from being inexplicable, that their occurrence might have been anticipated on our theory, and as fresh atolls are supposed to be in progressive formation by the subsidence of encircling barrier-reefs, a weighty objection, namely that the number of atolls must be increasing infinitely, might even have been raised, if proofs of the occasional destruction and loss of atolls could not have been adduced.
The disseverment of the larger Maldiva atolls.--The apparent progressive disseverment in the Maldiva Archipelago of large atolls into smaller ones, is, in many respects, an important consideration, and requires an explanation. The graduated series which marks, as I believe, this process, can be observed only in the northern half of the group, where the atolls have exceedingly imperfect margins, consisting of detached basin-formed reefs. The currents of the sea flow across these atolls, as I am informed by Captain Moresby, with considerable force, and drift the sediment from side to side during the monsoons, transporting much of it seaward; yet the currents sweep with greater force round their flanks. It is historically known that these atolls have long existed in their present state; and we can believe, that even during a very slow subsidence they might thus remain, the central expanse being kept at nearly its original depth by the accumulation of sediment. But in the action of such nicely balanced forces during a progressive subsidence (like that, to which by our theory this archipelago has been subjected), it would be strange if the currents of the sea should never make a direct passage across some one of the atolls, through the many wide breaches in their margins. If this were once effected, a deep-water channel would soon be formed by the removal of the finer sediment, and the check to its further accumulation; and the sides of the channel would be worn into a slope like that on the outer coasts, which are exposed to the same force of the currents. In fact, a channel precisely like that bifurcating one which divides Mahlos Mahdoo (Plate II., Figure 4.), would almost necessarily be formed. The scattered reefs situated near the borders of the new ocean-channel, from being favourably placed for the growth of coral, would, by their extension, tend to produce fresh margins to the dissevered portions; such a tendency is very evident (as may be seen in the large published chart) in the elongated reefs on the borders of the two channels intersecting Mahlos Mahdoo. Such channels would become deeper with continued subsidence, and probably from the reefs not growing up perpendicularly, somewhat broader. In this case, and more especially if the channels had been formed originally of considerable breadth, the dissevered portions would become perfect and distinct atolls, like Ari and Ross atolls (Plate II., Figure 6), or like the two Nillandoo atolls, which must be considered as distinct, although related in form and position, and separated from each other by channels, which though deep have been sounded. Further subsidence would render such channels unfathomable, and the dissevered portions would then resemble Phaleedoo and Moluque atolls, or Mahlos Mahdoo and Horsburgh atolls (Plate II., Figure 4), which are related to each other in no respect except in proximity and position. Hence, on the theory of subsidence, the disseverment of large atolls, which have imperfect margins (for otherwise their disseverment would be scarcely possible), and which are exposed to strong currents, is far from being an improbable event; and the several stages, from close relation to entire isolation in the atolls of the Maldiva Archipelago, are readily explicable.
We might go even further, and assert as not improbable, that the first formation of the Maldiva Archipelago was due to a barrier-reef, of nearly the same dimensions with that of New Caledonia (Plate II., Figure 5), for if, in imagination, we complete the subsidence of that great island, we might anticipate from the present broken condition of the northern portion of the reef, and from the almost entire absence of reefs on the eastern coast, that the barrier-reef after repeated subsidences, would become during its upward growth separated into distinct portions; and these portions would tend to assume an atoll-like structure, from the coral growing with vigour round their entire circumferences, when freely exposed to an open sea. As we have some large islands partly submerged with barrier-reefs marking their former limits, such as New Caledonia, so our theory makes it probable that there should be other large islands wholly submerged; and these, we may now infer, would be surmounted, not by one enormous atoll, but by several large elongated ones, like the atolls in the Maldiva group; and these again, during long periods of subsidence, would sometimes become dissevered into smaller atolls. I may add, that both in the Marshall and Caroline Archipelagoes, there are atolls standing close together, which have an evident relationship in form: we may suppose, in such cases, either that two or more encircled islands originally stood close together, and afforded bases for two or more atolls, or that one atoll has been dissevered. From the position, as well as form, of three atolls in the Caroline Archipelago (the Namourrek and Elato group), which are placed in an irregular circle, I am strongly tempted to believe that they have originated by the process of disseverment.
Irregularly formed Atolls.--In the Marshall group, Musquillo atoll consists of two loops united in one point; and Menchikoff atoll is formed of three loops, two of which (as may be seen in Figure 3, Plate II.) are connected by a mere ribbon-shaped reef, and the three together are sixty miles in length. In the Gilbert group some of the atolls have narrow strips of reef, like spurs, projecting from them. There occur also in parts of the open sea, a few linear and straight reefs, standing by themselves; and likewise some few reefs in the form of crescents, with their extremities more or less curled inwards. Now, the upward growth of a barrier-reef which fronted only one side of an island, or one side of an elongated island with its extremities (of which cases exist), would produce after the complete subsidence of the land, mere strips or crescent or hook-formed reefs: if the island thus partially fronted became divided during subsidence into two or more islands, these islands would be united together by linear reefs; and from the further growth of the coral along their shores together with subsidence, reefs of various forms might ultimately be produced, either atolls united together by linear reefs, or atolls with spurs projecting from them. Some, however, of the more simple forms above specified, might, as we have seen, be equally well produced by the coral perishing during subsidence on part of the circumference of an atoll, whilst on the other parts it continued to grow up till it reached the surface.
The Great Chagos Bank.--I have already shown that the submerged condition of the Great Chagos Bank (Plate II., Figure 1, with its section Figure 2), and of some other banks in the Chagos group, may in all probability be attributed to the coral having perished before or during the movements of subsidence, to which this whole area by our theory has been subjected. The external rim or upper ledge (shaded in the chart), consists of dead coral-rock thinly covered with sand; it lies at an average depth of between five and eight fathoms, and perfectly resembles in form the annular reef of an atoll. The banks of the second level, the boundaries of which are marked by dotted lines in the chart, lie from about fifteen to twenty fathoms beneath the surface; they are several miles broad, and terminate in a very steep slope round the central expanse. This central expanse I have already described, as consisting of a level muddy flat between thirty and forty fathoms deep. The banks of the second level, might at first sight be thought analogous to the internal step-like ledge of coral-rock which borders the lagoons of some atolls, but their much greater width, and their being formed of sand, are points of essential difference. On the eastern side of the atoll some of the banks are linear and parallel, resembling islets in a great river, and pointed directly towards a great breach on the opposite side of the atoll; these are best seen in the large published chart. I inferred from this circumstance, that strong currents sometimes set directly across this vast bank; and I have since heard from Captain Moresby that this is the case. I observed, also, that the channels or breaches through the rim, were all of the same depth as the central lagoon-like space into which they lead; whereas the channels into the other atolls of the Chagos group, and as I believe into most other large atolls, are not nearly as deep as their lagoons: for instance at Peros Banhos, the channels are only of the same depth, namely between ten and twenty fathoms, as the bottom of the lagoon for a space about a mile and a half in width round its shores, whilst the central expanse of the lagoon is from thirty-five to forty fathoms deep. Now, if an atoll during a gradual subsidence once became entirely submerged, like the Great Chagos Bank, and therefore no longer exposed to the surf, very little sediment could be formed from it; and consequently the channels leading into the lagoon from not being filled up with drifted sand and coral detritus, would continue increasing in depth, as the whole sank down. In this case, we might expect that the currents of the open sea, instead of any longer sweeping round the submarine flanks, would flow directly through the breaches across the lagoon, removing in their course the finer sediment, and preventing its further accumulation. We should then have the submerged reef forming an external and upper rim of rock, and beneath this portion of the sandy bottom of the old lagoon, intersected by deep-water channels or breaches, and thus formed into separate marginal banks; and these would be cut off by steep slopes, overhanging the central space, worn down by the passage of the oceanic currents.
By these means, I have scarcely any doubt that the Great Chagos Bank has originated,--a structure which at first appeared to me far more anomalous than any I had met with. The process of formation is nearly the same with that, by which Mahlos Mahdoo had been trisected; but in the Chagos Bank the channels of the oceanic currents entering at several different quarters, have united in a central space.
This great atoll-formed bank appears to be in an early stage of disseverment; should the work of subsidence go on, from the submerged and dead condition of the whole reef, and the imperfection of the south-east quarter a mere wreck would probably be left. The Pitt’s Bank, situated not far southward, appears to be precisely in this state; it consists of a moderately level, oblong bank of sand, lying from 10 to 20 fathoms beneath the surface, with two sides protected by a narrow ledge of rock which is submerged between 5 and 8 fathoms. A little further south, at about the same distance as the southern rim of the Great Chagos Bank is from the northern rim, there are two other small banks with from 10 to 20 fathoms on them; and not far eastward soundings were struck on a sandy bottom, with between 110 and 145 fathoms. The northern portion with its ledge-like margin, closely resembles any one segment of the Great Chagos Bank, between two of the deep-water channels, and the scattered banks, southward appear to be the last wrecks of less perfect portions.
I have examined with care the charts of the Indian and Pacific Oceans, and have now brought before the reader all the examples, which I have met with, of reefs differing from the type of the class to which they belong; and I think it has been satisfactorily shown, that they are all included in our theory, modified by occasional accidents which might have been anticipated as probable. In this course we have seen, that in the lapse of ages encircling barrier-reefs are occasionally converted into atolls, the name of atoll being properly applicable, at the moment when the last pinnacle of encircled land sinks beneath the surface of the sea. We have, also, seen that large atolls during the progressive subsidence of the areas in which they stand, sometimes become dissevered into smaller ones; at other times, the reef-building polypifers having entirely perished, atolls are converted into atoll-formed banks of dead rock; and these again through further subsidence and the accumulation of sediment modified by the force of the oceanic currents, pass into level banks with scarcely any distinguishing character. Thus may the history of an atoll be followed from its first origin, through the occasional accidents of its existence, to its destruction and final obliteration.
Objections to the theory of the formation of Atolls and Barrier-reefs.--The vast amount of subsidence, both horizontally or in area, and vertically or in depth, necessary to have submerged every mountain, even the highest, throughout the immense spaces of ocean interspersed with atolls, will probably strike most people as a formidable objection to my theory. But as continents, as large as the spaces supposed to have subsided, have been raised above the level of the sea,--as whole regions are now rising, for instance, in Scandinavia and South America,--and as no reason can be assigned, why subsidences should not have occurred in some parts of the earth’s crust on as great a scale both in extent and amount as those of elevation, objections of this nature strike me as of little force. The remarkable point is that movements to such an extent should have taken place within a period, during which the polypifers have continued adding matter on and above the same reefs. Another and less obvious objection to the theory will perhaps be advanced from the circumstance, of the lagoons within atolls and within barrier-reefs never having become in any one instance during prolonged subsidences of a greater depth than sixty fathoms, and seldom more than forty fathoms; but we already admit, if the theory be worth considering, that the rate of subsidence has not exceeded that of the upward growth of the coral on the exterior margin; we are, therefore, only further required to admit, that the subsidence has not exceeded in rate the filling up of the interior spaces by the growth of the corals living there, and by the accumulation of sediment. As this filling up must take place very slowly within barrier-reefs lying far from the land, and within atolls which are of large dimensions and which have open lagoons with very few reefs, we are led to conclude that the subsidence thus counter-balanced, must have been slow in an extraordinary degree; a conclusion which accords with our only means, namely, with what is known of the rate and manner of recent elevatory movements, of judging by analogy what is the probable rate of subsidence.
In this chapter it has, I think, been shown, that the theory of subsidence, which we were compelled to receive from the necessity of giving to the corals, in certain large areas, foundations at the requisite depth, explains both the normal structure and the less regular forms of those two great classes of reefs, which have justly excited the astonishment of all persons who have sailed through the Pacific and Indian Oceans. But further to test the truth of the theory, a crowd of questions will occur to the reader: Do the different kinds of reefs, which have been produced by the same kind of movement, generally lie within the same areas? What is their relation of form and position,--for instance, do adjoining groups of atolls, and the separate atolls in these groups, bear the same relation to each other which islands do in common archipelagoes? Have we reason to believe, that where there are fringing-reefs, there has not lately been subsidence; or, for it is almost our only way of ascertaining this point, are there frequently proofs of recent elevation? Can we by this means account for the presence of certain classes of reefs in some large areas, and their entire absence in others? Do the areas which have subsided, as indicated by the presence of atolls and barrier-reefs, and the areas which have remained stationary or have been upraised, as shown by fringing-reefs, bear any determinate relation to each other; and are the dimensions of these areas such as harmonise with the greatness of the subterranean changes, which, it must be supposed, have lately taken place beneath them? Is there any connection between the movements thus indicated, and recent volcanic action? All these questions ought to receive answers in accordance with the theory; and if this can be satisfactorily shown, not only is the theory confirmed, but as deductions, the answers are in themselves important. Under this latter point of view, these questions will be chiefly considered in the following chapter.
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End of part 9 of 14
Continue readingChapter VI: On the Distribution of Coral-Reefs with Reference to the Theory of Their Formation (Part 1)
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