Galapagos finches with different shaped beaks perched near a giant tortoise on a black volcanic shore, with a three masted sailing ship anchored behind them.

Starvard University · Evolution

Evolution 101 with Darwin

Sail with a young naturalist, dig up giant extinct beasts, and watch one careful idea grow: natural selection, explained from Darwin's own books.

Finn

Hello, my name is Finn, and I'm going to help you understand this course.

Here it is, explained like you are 10:

Imagine you are twenty two, and you get a job sailing around the world for almost five years, collecting rocks, fossils, and animals. That is what happened to Charles Darwin. He came home with boxes of specimens and one huge question: why do living things fit their homes so well, and why do they differ a little from island to island? In this course you follow his clues in order: giant fossil bones, island birds, tame pigeons, and too many baby elephants. By the end you will understand his answer, natural selection, and also the hard parts he was honest about.

The whole course in brief

What you will be able to do after this course

  1. Tell the story of the Beagle voyage and what Darwin actually did on it.
  2. Explain what South American fossils showed about past and present life.
  3. Describe how island birds raised the question of species changing.
  4. Use Darwin's pigeons to explain variation and selection by breeders.
  5. State the struggle for existence in plain words, with his elephant example.
  6. Define natural selection the way Darwin defined it himself.
  7. List the difficulties Darwin admitted, and how he answered them.
  8. Explain Darwin's place in science today, fairly and without caricature.

The seven lessons

Lesson 1

A young naturalist sets sail

Charles Darwin was in his early twenties when his Cambridge teacher, the Reverend Professor Henslow, helped him toward the chance of a lifetime. HMS Beagle, a ten gun brig under Captain Fitz Roy, sailed from Devonport on 27 December 1831. The expedition was to finish surveying Patagonia and Tierra del Fuego, survey the coasts of Chile and Peru and some Pacific islands, and carry a chain of timing measurements around the world. Darwin went as the ship's naturalist.

His real work was patient collecting. He rode inland, dug in cliffs, shot and skinned birds, bottled insects, and shipped crates of specimens home to Henslow, who cared for them during his absence. Darwin later wrote that two kinds of facts struck him most: how the inhabitants of South America were distributed, and how the living animals of that continent related to its past inhabitants. Those two puzzles, distribution and fossils, became the seed of everything.

The voyage lasted almost five years. Darwin came home not with a finished theory but with notebooks, specimens, and a question he would spend two decades answering.

A collector first. Darwin's theory began as crates of real specimens and two puzzles: where species live, and how living forms relate to fossil ones.

Lesson 2

Fossils, earthquakes, and the Galápagos

At Punta Alta in Argentina, Darwin dug the bones of nine great extinct quadrupeds from a space about two hundred yards square. There were giant ground sloth relatives such as Megatherium and Mylodon, a Scelidotherium as large as a rhinoceros, an animal in armadillo like armour, an extinct horse, and the Toxodon, which he called perhaps one of the strangest animals ever discovered: elephant sized, yet with teeth linking it to the gnawing animals. Living sloths, armadillos, and anteaters in the same land looked like smaller relatives of these giants, a pattern he never forgot.

In Chile he felt the great earthquake of 1835, which overthrew Concepcion, and saw land raised by such shocks. Slow rises and sinkings, repeated over ages, could build mountains and, he later argued in his book on coral reefs, grow atolls as volcanic land gradually sank beneath the sea.

Then came the Galapagos. Years later, comparing birds from the separate islands with each other and with mainland birds, Darwin wrote that he was struck by how vague and arbitrary the line between species and varieties seemed. Island forms were similar, yet not the same.

Patterns, not curiosities. Fossils resembled living neighbours, earthquakes showed land moving, and island birds differed slightly from place to place.

Lesson 3

Variation under domestication

Darwin opened his big book not in the wild but in the farmyard. His reasoning was simple: before asking how species change in nature, study how much living things can change in the hands of people. So he kept pigeons. He bought every breed he could, joined two London pigeon clubs, and received skins from India and Persia. What he found astonished him, and he called the diversity of the breeds something astonishing.

Compare the English carrier, with its long beak and wattled skin, and the short faced tumbler, whose beak is almost like a finch's. The fantail carries thirty or even forty tail feathers where twelve or fourteen is normal for the family. The pouter inflates an enormous crop. Darwin reckoned at least a score of pigeons could be shown to a bird expert, passed off as wild birds, and ranked as separate species.

Yet he concluded that all of them descended from one ancestor, the rock pigeon, Columba livia. Breeders had built the differences by keeping, generation after generation, the slight variations they liked. Selection, patiently applied to small differences, clearly had great power.

Pigeons prove change is possible. One ancestor, the rock pigeon, became carriers, tumblers, and fantails because breeders kept choosing slight differences.

Lesson 4

The struggle for existence

Darwin's next step was arithmetic. Every living thing tends to increase in geometrical ratio, producing far more offspring than the world can hold. His calm example was the slowest breeder he knew, the elephant. Assume a pair breeds from age thirty to ninety and raises three pairs of young, and within five centuries, he calculated, fifteen million elephants would be alive, descended from that first pair. Since the world is plainly not buried in elephants, most offspring must die before breeding.

Darwin said plainly where this idea came from: it is the doctrine of Malthus, the writer on human population, applied with manifold force to the whole animal and vegetable kingdoms. In nature there is no artificial increase of food to rescue the surplus.

He used the phrase struggle for existence in a wide, metaphorical sense. It includes a plant at the desert's edge depending on moisture, seedlings competing for light, and one animal eating another. The point is not constant fighting. It is that more are born than can possibly survive, so even a slight advantage matters enormously.

Too many mouths, too little room. Because every species produces more young than can survive, any small helpful difference can decide who leaves descendants.

Lesson 5

Natural selection

Now the pieces join. Living things vary, and much of that variation is inherited. More individuals are born than can survive. So Darwin asked: if breeders, choosing for a few generations, can reshape a pigeon, what happens in nature over thousands of generations, where the choice is made by survival itself?

His answer is worth hearing in his own words. Individuals having any advantage, however slight, over others would have the best chance of surviving and of procreating their kind, while any variation in the least degree injurious would be rigidly destroyed. Then his definition, which this course has been building toward: this preservation of favourable variations and the rejection of injurious variations, I call Natural Selection.

Notice what the definition does not say. It does not say the strongest wins, or that animals try to evolve, or that nature has a plan. It says that helpful differences tend to be kept, harmful ones tend to be lost, and that tiny steps, added up over immense time, can produce large change, just as the breeder's small choices produced the fantail.

Kept or lost, step by step. Natural selection is simply this: favourable variations tend to be preserved, injurious ones tend to be destroyed.

Lesson 6

Difficulties Darwin admitted

Darwin devoted whole chapters of his book to the objections against his own theory, and he did not choose soft ones. The eye was his most famous case. To suppose that the eye, with all its contrivances for adjusting focus and admitting light, could have been formed by natural selection seemed, he wrote, absurd in the highest possible degree. His reply was to look for gradations: living animals show every step from a light sensitive nerve to a complex eye, and if each step is useful, the difficulty lies in our imagination, not in nature.

Other admissions were just as frank. Neuter insects, the sterile worker ants and bees that cannot pass on their own traits, he called one of the gravest difficulties, and answered it by arguing that selection can act on the family, not only the individual. The fossil record troubled him too: he named a chapter on its imperfection, arguing that the earth's crust is a vast museum whose collections were made only at long intervals, so most transitional forms were never preserved.

He published these difficulties himself, in his own book, for every critic to read.

He listed his own problems. The eye, sterile insects, and the patchy fossil record were difficulties Darwin raised himself and answered as honestly as he could.

Lesson 7

Darwin's place in science today, and how people of faith have engaged him

Darwin's book appeared in 1859, more than twenty years after he first sketched the idea, and only when Alfred Russel Wallace reached nearly the same conclusion independently and sent him a memoir. Darwin's friends Lyell and Hooker arranged for both men's work to be presented together, a fair settlement. Since then, evolution by natural selection has become the organising idea of biology, extended by genetics, a science Darwin never knew, which supplied the mechanism of inheritance his theory needed.

People of faith have engaged Darwin in many honest ways. Some of his earliest defenders were churchmen and believers, who argued that discovering how life develops is not the same as explaining it away, and many scientists and theologians since have held their faith and accepted evolution together, seeing no necessary conflict. Others have disagreed, for reasons they take seriously. The arguments are best read in that same fair spirit.

Darwin himself drifted away from the Christian faith of his youth, a loss he connected in his letters and memoir with grief, including the death of his beloved daughter Annie, and with long doubt. He was no enemy of religion, and when he died in 1882 he was buried in Westminster Abbey, close to Newton, an honour from his own country.

Read him fairly. Darwin changed biology, believers have both defended and debated him in good faith, and he deserves to be met honestly, not as a caricature.

Sources

The Books Behind This Course

This course is taught from classic texts that are free for anyone to read. Every method in the lessons traces back to one of these books.

Graduation

You have read Darwin, not just about him

You started on a ship's deck in 1831 and finished with one of the most tested ideas in science. Keep Darwin's own habits as you go: collect real facts, reason slowly, state the objections against yourself, and stay fair to people who see things differently. That is good science, and it is good manners too. Congratulations on finishing Evolution 101 with Darwin.

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